A possible Pararcus diepenbroeki vertebra from the Vossenveld Formation (Triassic, Anisian), Winterswijk, the Netherlands

Abstract

An isolated, completely ossified vertebra tentatively ascribed to the non-cyamodontid placodont Pararcus diepenbroeki is described from the Anisian Vossenveld Formation in Winterswijk, the Netherlands, and compared to other material from the same locality. This fossil is the first completely ossified vertebra of the taxon and most likely originates from an adult specimen. It was recovered c. 16 m deeper in the stratigraphy than previously described material of the species, which is thus far known only from Winterswijk. Based on the slanting angle of the transverse process, the vertebra is interpreted to originate from the dorsal region. Besides the overall agreements in morphology that warrant a tentative identification as Pararcus diepenbroeki, the newly described vertebra deviates from other known Pararcus vertebrae in the presence of a longer, well-ossified neural spine and a strongly constricted, less pachyostotic and ovaloid vertebral centrum. General agreement in morphology with previously described vertebrae suggests this novel condition indicates a different anatomical position and perhaps a varied ossification pattern.